It seems the anode is the screen grid. And the anode is used as an anode of a diode to crate to side chain DC signal for gain control or reduction?!
US2373180
https://worldwide.espacenet.com/publicationDetails/originalDocument?CC=US&NR=2373180A&KC=A&FT=D&ND=3&date=19450410&DB=EPODOC&locale=en_EP
http://euroelectron.blogspot.com/2018/08/interesting-automatic-gain-control.html
See also how it may be done automaticly by the diode rectification of the grid:
The Neumann U47 compressing - euroelectron
Operating Features of the Audion - E. H. Armstrong
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